Historical overview of mugilid systematics, with description of Paramugil (Teleostei; Mugiliformes; Mugilidae), new genus

by Javad Ghasemzadeh, Walter Ivantsoff  and Aarn

aqua, International Journal of Ichthyology, pp. 9-22, Volume 8, Issue 1 – February 2004


The history of the systematic relationships of the mugilids is reviewed, concluding with the modern concept of Mugilidae comprising 17 genera with 80 species, one of five taxa comprising ‘Smegmamorpha’. Paramugil, new genus, is erected for P. georgii and P. parmata, and 18 diagnostic morphological and osteological differences between Paramugil and Liza and/or Mugil and/or Valamugil listed.

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Anderson, M. 1982. The identification of the British grey mullets. Journal of Fish Biology, 20: 33-38. Athanassopoulos, G. 1919. Contributo alla distinzione delle specie Mediterranee del genere Mugil. Annali del Museo Civico di Storia Naturale (di Genova) Giacomo Doria, 3 (8): 254-269.

Autem, M. & Bonhomme. 1980. Éléments de systematique biochimique chez les Mugilides de Mediterranée. Biochemical and Systematic Ecology, 8: 305-308.

Berg, L. S. 1940. Classification of Fishes both Recent and Fossil. Tr. Zool. Inst. Akad. Nauk SSSR, 5 (2) (In Russian).

Berra, T. M. 1981. An Atlas of Distribution of the Freshwater Fish Families of the World. University of Nebraska Press, London, 197 pp.

Bleeker, P. 1852a. Zesde bijdrag tot de kennis der ichthyologische fauna van Borneo. Visschen van Pamangkat, Bandjermassing, Braboekarta en Sampit. Natuurkundik Tijdshjrift voor Nederlandsh Indië. (Batavia), 3: 407-442.

Bleeker, P. 1852b. Nieuwe bijdrag tot de kennis der ichthyologische fauna van Ceram. Natuurkundik Tijdshjrift voor Nederlandsh Indië. (Batavia), 3: 689-714.

Bleeker, P. 1855. Verslag van eenige vischverzamelingen van visschen van Oost-Java. Natuurkundik Tijdshjrift voor Nederlandsh Indië. (Batavia), 9: 391-414.

Bleeker, P. 1859a. Conspectus specierum Mugilis Archipelagi indici analyticus. Natuurkundik Tijdshjrift voor Nederlandsh Indië. (Batavia), 16: 275-81.

Bleeker, P. 1859 b. Derde bijdrag tot de kennis der vischfauna van Soembawa. Natuurkundik Tijdshjrift voor Nederlandsh Indië. (Batavia), 19: 434-440.

Bleeker, P. 1860. Diertiende bijdrag tot de kennis der vischfauna van Borneo. Acta Societatis Scientiarum Indo-Neérlandica (Batavia), 8: 1-64.

Bleeker, P. 1861. Iets over de vischfauna van heteiland Pinang. Verslagen. Academie van. Amsterdam, 12: 64-80.

Borcea, I. 1934. Données pour servir à la systématique et à la biologie des Mugilides Formes de la Mer Noire (litoral Roumain). Annales des Sciences de l’Université. De Jassy, 19: 247-286.

Cadenat, J. 1954. Notes d’ichthyologie ouestafricaine VIII. Sur les mulets de la côte occidentale d’Afrique. Bulletin de l’Institut Francais d’Afrique Noire, 16: 584-591.

Cambrony, M. 1984. Identification et périodicité du recrutement des juvéniles de Mugilidae dans les étangs littoraux du Languedoc-Roussillon. Vie Milieu, 34: 221-227.

Campton, D. E. & B. Mahmoudi. 1991. Allozyme variation and population structure of striped mullet (Mugil cephalus) in Florida. Copeia, 1991 (2): 485-492.

Cantor, T. E. 1850. Catalogue of Malayan Fishes. Journal of the Asiatic Society of Bengal, 18 (2): 983-1043.

Chervinski, J. 1984. Using scales for identification of four Mugilidae species. Aquaculture, 38: 79-81.

Chubb, C. F., Potter, I. C., Grant, C. J., Lenanton, R. C. J. & J. Wallace. 1981. Age structure, growth rates and movement of sea mullet Mugil cephalus L., and yellow eyemullet, Aldrichetta forsteri (Valenciennes) in the Swan Avon River system, Western Australia. Australian Journal of Marine and Freshwater Research, 32 (4): 605-628.

Cockerell, T. D. A. 1913. The scales of some Queensland fishes. Memoirs of the Queensland Museum, 2: 51-9.

Crosetti, D., Avise, J. C., Placidi, F., Rossi, A. R. & L. Sola. 1993. Geographic variability in the grey mullet Mugil cephalus: preliminary results of mt DNA and chromosome analyses. Aquaculture, 111: 95-101.

Crosetti, D., Nelson, W. S. & J. C. Avise. 1994. Pronounced genetic structure of mitochondrial DNA among populations of the circumglobally distributed grey mullet (Mugil cephalus). Journal of Fish Biology, 44: 47-58.

Day, F. 1865a. On the Fishes of Cochin, on the Malabar coast of India. Pt 1. Acanthopterygii. Proceedings of the Zoological Society of London, 1865: 2-40.

Day, F. 1865b. The Fishes of Malabar. Bernard Quaritch, London, 293 pp.

Day, F. 1878. The Fishes of India, being a Natural History of the Fishes known to inhabit the Seas and Fresh waters of India, Burma, and Ceylon (vol. 1). Today and Tomorrow’s Book Agency, New Delhi, 816 pp.

Day, F. 1889. The Fauna of British India, including Ceylon and Burma. Fishes (vol 2). Taylor and Francis, London, 509 pp.

Deva-Sundaram, M. P. 1951. Systematics of Chilka Mullets with a key to their Identification. Journal of the Zoological Society of India, 3 (1): 19-25.

Ebeling, A. W. 1957. The dentition of eastern Pacific mullets, with special reference to adaptation and taxonomy. Copeia, 1957 (3): 173-185.

Ebeling, A. W. 1961. Mugil galapagensis, a new mullet from the Galapagos Islands, with notes on related species and a key to the Mugilidae of the eastern Pacific. Copeia, 1961: 295-304.

Farrugio, H. 1977. Clés commentées pour la détermination des adults et des alevims de Mugilidae de Tunisie. Cybium, 3rd Ser. (2): 57-73.

Fowler, H. W. 1925. Fishes from Natal, Zululand, and Portuguese East Africa. Proceedings of the Academy of Natural Sciences of Philadelphia, 77: 187-268.

Fowler, H. W. 1928. The Fishes of Oceania. Memoirs of the Bernice P. Bishop Museum, 10: 1-546.

Fowler, H. W. 1931. The fishes of Oceania. Supplement 1. Bernice P. Bishop Museum- Memoir, 10 (5): 313-81.

Fowler, H. W. 1934. The fishes of Oceania. Memoirs of the Bernice P. Bishop Museum Supplement, 2: 11 (6): 385-466.

Fowler, H. W. 1935. Zoological Results of the Third de Schauensee Siamese Expedition. Part 8- Fishes Obtained in 1936. Proceedings of the Academy of Natural Sciences of Philadelphia, 139: 125-264.

Fowler, H. W. 1936. The marine fishes of West Africa. Bulletin of the American Museum of Natural History, 70 (1): 1-605.

Fowler, H. W. 1939. Zoological Results of the Third de Schauensee Siamese Expedition, pt. IX. Additional fishes obtained in 1936. Proceedings of the Academy of Natural Sciences of Philadelphia, 91: 39-76.

Fowler, H. W. 1959. Fishes of Fiji. Government of Fiji. Suva, 670 pp.

Ghasemzadeh, J. 1998. Phylogeny and Systematics of Indo-Pacific mullets (Teleostei: Mugilidae) with special reference to the mullets of Australia. Unpublished Ph.D Thesis, Macquarie University, Sydney, 397 pp.

Gosline, W. A. 1968. The suborders of perciformfishes. Proceedings of the United States National Museum, 124: 1-78.

Gosline, W. A. 1971. Functional morphology and classification of teleostean fishes. University of Hawaii Press, Honolulu. 208 pp.

Grant, C. J. & A. V. Spain. 1975a. Reproduction growth and size allometry of Mugil cephalus Linnaeus (Pisces: Mugilidae) from North Queensland inshore waters. Australian Journal of Zoology, 23: 181-201.

Grant, C. J. & A. V. Spain. 1975b. Reproduction growth and size allometry of Valamugil seheli (Forskål) (Pisces: Mugilidae) from North Queensland inshore waters. Australian Journal of Zoology, 23: 463-74.

Grant, C. J. & A. V. Spain. 1975c. Reproduction growth and size allometry of Liza vaigiensis (Quoy and Gaimard) (Pisces: Mugilidae) from North Queensland inshore waters. Australian Journal of Zoology, 23: 475-85.

Grant, C. J. & A. V. Spain. 1977. Variation of the body shape of three species of Australian Mullets (Pisces: Mugilidae) during the course of development. Australian Journal of Marine and Freshwater Research, 28 (6): 723- 738.

Grant, C. J., Spain, A. V. & P. N. Jones. 1977. Studies of sexual dimorphism and other variations in nine species of Australian Mullets (Pisces: Mugilidae). Australian Journal of Zoology, 25 (4): 615-630.

Grant, E. M. 1965. Guide to Fishes. The Department of Harbours and Marine, Brisbane, 896 pp.

Grant, E. M. 1987. Fishes of Australia. E. M. Grant Ltd., Brisbane, 1016 pp.

Greenwood, P. H., Rosen, D. E., Weitzman, S. H. & G. S. Myers. 1966. Phyletic studies of teleostean fishes with a provisional classification of living forms. Bulletin of the American Museum of Natural History, 131 (4): 339-456.

Günther, A. 1861. Catalogue of the acanthopterygian fishes in the British Museum (vol. 3). Trustees of the British Museum, London, 586 pp.

Gunther, G., Sulya, L. L. & B. E. Box. 1961. Some evolutionary patterns in fishes’ blood. Biological Bulletin, 121 (2): 302-6.

Harris, J. A. 1968. The yellow-eyemullet. Age structure, growth rate and spawning cycle of a population of yellow- eye mullet Aldrichetta forsteri (Cuv.and Val.) from the Coorong Lagoon, South Australia. Transactions of the Royal Society of South Australia, 92: 37-50.

Harrison, I. J. & G. J. Howes. 1991. The pharyngobranchial organ of mugilid fishes; its structure, variability, ontogeny, possible function and taxonomic utility. Bulletin of the British Museum of Natural History (Zoology), 57 (2): 111-132.

Harrison, I. J. & H. Senou. (in press). Mugilidae. In: Carpenter, C., & Niem, V. H. (eds), FAO species identification guide for fishery purposes. The living marine resources of theWestern Central Pacific. FAO, Rome.

Herre, A. W. 1936. Fishes of the Crane Pacific Expedition. Publications of the Field Museum of Natural History 353, Zoology Series, 21: 1-472.

Herre, A. W. 1953. Checklist of Philippine Fishes. Research Report 20. Fish and Wildlife Service, United States Government Printing Office, 975 pp.

Hollister, G. 1937. Caudal skeleton of Bermuda shallow water fishes. II. Order Percamorphi, suborder Percesoces: Atherinidae, Mugilidae, Sphyraenidae. Zoologica, 22 (3): 265-279.

Hotta, H. 1955. On the mature mugilid fish from Kabashima, Nagasaki Pref., Japan, with additional notes on the intestinal convolution of Mugilidae. Japanese Journal of Ichthyology, 4: 162-169.

Hotta, H. & I. S. Tung. 1966. Identification of fishes of the family Mugilidae based on the pyloric caeca and the position of inserted first interneural spine. Japanese Journal of Ichthyology, 14: 62-66.

Ingham, S. E. 1952. The biology and taxonomy of the Mugilidae. Unpublished MS thesis, British Museum (Natural History), London, 276 pp.

Ishiyama, R. 1951. Revision of the Japanese mugilid fishes, especially based upon the osteological characters of the cranium. Japanese Journal of Ichthyology, 1 (4): 238-250.

Jacot, A. P. 1920. Age, growth and scale characters of the mullet Mugil cephalus and Mugil curema. Transactions of the AmericanMicroscopy Society, 39: 199-229.

Johnson, G. D. & C. Patterson. 1993. Percomorph phylogeny: a survey of acanthomorphs and a new proposal. Bulletin of Marine Science, 52 (1): 554-626.

Jordan, D. S. & B. W. Evermann. 1896a. The Fishes of North and Middle America: A descriptive catalogue of the species of fish-like vertebrates found in the waters of America, north of the isthmus of Panama. Part 1. Bulletin of the United States National Museum, 47: 1-1240.

Jordan, D. S. & B.W. Evermann. 1896a. A checklist of fishes and fish like vertebrates of North and Middle America. Report of the United States Fisheries Commission, 21: 207-584.

Jordan, D. S. & R. E Richardson. 1908. Fishes from the Islands of the Philippine Archipelago. Bulletin of the United States Bureau of Fisheries, 27: 233-87.

Jordan, D. S. & J. Swain. 1884. A review of the American species of marine Mugilidae. Proceedings of the United States National Museum, 7: 261-275.

Karunasinghe, W. P. N. 1995. Population Variability and inter-relationship of five species of mullet (Pisces: Mugilidae). Unpublished Ph D Thesis, Macquarie University, Sydney, 160 pp.

Kobelkowsky, A. D. & Resendez. 1972. Estudio comparativo del endosqueleyo de Mugil cephalus y Mugil curema (Pisces, Perciformes). Anales del Instituto Ciencias del Mar y Limnologia, Universidad Nacional Autonoma de Mexico, 43 (1): 33-81.

Lake, J. S. 1971. Freshwater Fishes and Rivers of Australia. Thomas Nelson and Sons, Melbourne, 61 pp. Langi, S. A., Latu T. F. & S. Tulua. 1990. Preliminary study of the biology of mullets (Pisces: Mugilidae) from Nuku’ Alofa, Tonga. Papers on Fisheries Science in Pacific Islands, 1: 37-42.

Lee, C. L., & D. S. Joo. 1994. Synopsis of family Mugilidae (Perciformes) from Korea. Bulletin of the Korean Fisheries Society, 27 (6): 814-824.

Lee, S. C., Chang, J. T. & Y. Y. Tsu. 1995. Genetic relationships of four Taiwan mullets (Pisces: Perciformes: Mugilidae). Journal of Fish Biology, 46: 159-162.

Lee, S. C., Chang, H. L. & J. T. Chang. 1996. Allozyme variation in the large scale mullet Liza macrolepis (Perciformes: Mugilidae) from coastal waters of western Taiwan. Zoological Studies, 35 (2): 85-92.

Lenanton, R. C. J., Potter, I. C., Lonergan, N. R., & P. J. Chrystal. 1984. Age structure and changes in abundance of three important species of teleost in a euphrotic estuary (Pisces: Teleostei). Journal of Zoology, London, 203: 311-327.

Leviton, A. E., Robert, H., Gibbs, JR., Heal, E. & C. E. Dawson. 1985. Standards in Herpetology and Ichthyology: Part I. Standard Symbolic Codes for Institutional Resource Collections in Herpetology and Ichthyology. Copeia, 1985 (3): 802-832.

Liu, C. H. & S. C. Shen. 1991a. A revision of the mugilid fishes from Taiwan. Bulletin of the Institute of Zoology Academia Sinica, 30 (4): 273-288.

Liu, C. H. & S. C. Shen. 1991b. Lepidology of the Mugilid fishes. Journal of the Taiwan Museum, 44 (2): 321-357.

Luther, G. 1977. New characters for consideration in the taxonomic appraisal of grey mullets. Journal of the Marine Biology Association of India, 19 (1): 1-9.

Macleay, W. 1880. On the Mugilidae of Australia. Proceedings of the Linnaean Society of New South Wales, 4: 410-27.

Macleay, W. 1884a. On a new and remarkable fish of the family Mugilidae from the interior of New Guinea. Proceedings of the Linnaean Society of New South Wales, 8 (1): 2-6.

Macleay, W. 1884b. Notes on the collection of fishes from the Burdekin and Mary Rivers, Queensland. Proceedings of the Linnaean Society of New South Wales, 8 (2): 199-212.

Marshall, T. C. 1964. Fishes of the Great Barrier Reef and coastal waters of Queensland. Angus and Robertson, Sydney, 566 pp.

Marshall, T. C. 1982. Tropical Fishes of the Great Barrier Reef. Angus and Robertson, Sydney, 239 pp.

Matsubara, K. 1955. Fish morphology and hierarchy. Part I (Ishizuki-Shoten: Tokyo) (in Japanese), pp. 486-492.

McCulloch, A. R. 1921a. Check List of The Fishes and Fish-Like Animals of New South Wales. II. Australian Zoology, 2 (2): 24-68.

McCulloch, A. R. 1921b. Studies in Australian Fishes. 7. Records of the Australian Museum, 13: 123-142.

McCulloch, A. R. 1922. The Fishes of New South Wales. Australian Zoology Handbook (first ed). McCulloch, A. R. 1927. The Fishes and Fish-Like Animals of New South Wales. Australian Zoology Handbook (second ed).

McCulloch, A. R. 1929. A Check List of The Fishes Recorded from Australia. Memoirs of the Australian Museum, 5: 1-534.

McCulloch, A. R. 1934. The Fishes and Fish-Like Animals of New South Wales (third ed). Royal Zoological Society of New South Wales, Sydney, 104 pp.

McCulloch, A. R. & G. P. Whitley. 1925. Fishes recorded from Queensland. Memoirs of the Queensland Museum, 8 (2): 125-82.

Menezes, M. R., Martins, M. & S. Naik. 1992. Interspecific genetic divergence in grey mullets from the Goa region. Aquaculture, 105: 117-129.

Merrick, J. R. & G. E. Schmida. 1984. Australian Freshwater Fishes. Biology and Management. Griffin Press, Sydney, 409 pp.

Mohr, E. 1927. Mugiliden-Studien. Zoologisches Jahrbuch, 54: 177-20.

Mohsin, A. K. M. 1978. The osteology of the grey mullet, Liza subviridis (Valenciennes). The branchiocranium, vertebral column, and girdles. Malayan Nature Journal, 31 (4): 202-218.

Morovic, D. 1953. Sur la détermination des muges adriatiques d’après la forme de l’otolith sagitta. Notes Inst. Oceanogr. Split, 9: 1-7.

Munro, I. S. R. 1955. Marine and Fresh water Fishes of Ceylon. Department of External Affairs, Canberra, 351 pp.

Munro, I. S. R. 1956. Handbook of Australian Fishes. CSIRO publications, Canberra, 172 pp.

Munro, I. S. R. 1967. The Fishes of New Guinea. Department of Agriculture, Stock and Fisheries, Port Moresby, 650 pp.

Nelson, J. S. 1984. Fishes of the world (second ed). John Wiley and Sons, New York, 523 pp.

Nelson, J. S. 1994. Fishes of the world (third ed). John Wiley and Sons, New York, 600 pp.

Ogilby, J. D. 1887. On a new genus and species of Australian Mugilidae. Proceedings of the Zoological Society of London, 1887: 614-616.

Ogilby, J. D. 1897. New genera and species of Australian fishes. Proceedings of the Linnaean Society of New South Wales, 22: 62-95.

Ogilby, J. D. 1908. New or little known fishes in the Queensland Museum. Annals of the Queensland Museum, 9: 1-41.

Ogilby, J. D. 1915. The Commercial Fishes and Fisheries of Queensland. Queensland Government Printer, Brisbane.

Oren, O. H., 1981. Aquaculture of grey mullets. Cambridge University Press, London, 507 pp.

Oshima, M. 1922. A review of the fishes of the family Mugilidae found in the waters of Formosa. Annals of the Carnegie Museum, 13: 240-59.

Pandy, A. K. & G. S. Sandhu. 1992. Encyclopaedia of fishes and fisheries of India (vol. VI). Anmol Publications. New Delhi.

Parenti, L. R. 1993. Relationships of atherinomorph fishes (Teleostei). Bulletin of Marine Science, 52: 170-196.

Perlmutter, A., Bogard, L. & Pruginin 1957. Use of the estuarine and sea fish of the family Mugilidae (grey the Fisheries Council of the Mediterranean FAO, 4: 289-304.

Peterson, G. L. & Z. H. Shehadeh. 1975. Subpopulation of the Hawaiian striped mullet Mugil cephalus: analysis of variations of nuclear eye-lens protein electropherograms and nuclear eye-lens weights. Marine Biology, 11: 52-60.

Pillay, S. R. 1972. A bibliography of the grey mullets. FAO Fisheries Technical Paper Number 109, Rome, 99 pp.

Pillay. T. V. R. 1951. Structure and development of the scales of five species of grey mullets of Bengal. Proceedings of the National Institute of Science, India, 17 (6): 413-24.

Pillay. T. V. R. 1962. A revision of Indian Mugilidae. Journal of the Bombay Natural History Society, 59: 254-270.

Poll, M. 1959. Poissons V. Téléostéens Acanthopterygiens (deuxième partie) Résultats Scientifiques. Expédition for Océanographie Belge dans les Eaux Côtières Africaines de l’Atlantique Sud, 4 (3B): 1-417.

Popov, A. M. 1929. A preliminary revision of the Russian Mullets (Pisces: Mugilidae). Comptes Rendues of the Academy of Sciences of the USSR, 1929 (A): 243-247.

Popov, A. M. 1930. Mullets of Europe (Mugilidae) with description of a new species from the Pacific Ocean. Trudi Sevastopol’skoi Biologicheskoi Stantsii, 2: 47-125 (in Russian, English summary).

Popov, A. M. 1931. Some remarks on the genera of the family Mugilidae (Pisces) Annals of the Museum of Zoology of the Academy of Sciences of the USSR, 32: 117-125.

Reay, P. J. & V. Cornell. 1988. Identification of grey mullet (Teleostei: Mugilidae) juveniles from British waters. Journal of Fish Biology, 32: 95-99.

Rosen, D. E. 1964. The relationships and taxonomic position of the half-beaks, killifishes, silversides and their relatives. Bulletin of the American Museum of Natural History, 127 (5): 219-267.

Rossi, A. R., Crosetti, D., Gornung, E. & L. Sola. 1996. Cytogenetic analysis of global populations of Mugil cephalus (striped mullet) by different staining techniques and fluorescent in situ hybridization. Heredity, 76: 77-82.

Rossi, A. R., Capula, D., Crosetti, D., Campion, E. & L. Sola, 1998. Genetic divergence and phylogenetic inferences in five species of Mugilidae (Pisces: Perciformes). Marine Biology, 131: 213-218.

Roughley, T. C. 1964. Fish and Fisheries of Australia. Angus and Robertson, Sydney, 343 pp. Roxas, H. A. 1934. A review of Philippine Mugilidae. Philippine Journal of Science, 54: 393-431.

Roxas, H. A. & C. Martin. 1937. A check list of Philippine fishes. Department of Agriculture and Commerce, Manilla, 314 pp.

Schultz, L. P. 1946. A revision of the genera of mullets, fishes of the family Mugilidae, with descriptions of three new genera. Proceedings of the United States National Museum, 96: 377-395.

Schultz, L. P. 1953. Family Mugilidae. In: L. P. Schultz, E. S. Herald, E. A. Lachner, A. D. Welander and L. P. Woods. Fishes of the Marshal and Marianas Islands. Vol. 1. Bull. U.S. Nat. Mus., 202 (1): 310-22., pls. 23-24.

Senou, H. 1988. Phylogenetic Interrelationships of the Mullets (Pisces: Mugilidae). Unpublished PhD thesis, Tokyo University, 172 pp (in Japanese).

Serventi, M., Harrison, I. J., Torriecelli & G. Gandolfi. 1996. The use of pigmentation and morphological characters to identify Italian mullet fry. Journal of Fish Biology, 49 (6): 1163- 1173.

Shehadeh, Z. H., Kuo, C. M. & K. K. Milisen. 1973. Induced spawning of grey mullet M. cephalus L. with fractionated salmon pituitary extract. Journal of Fish Biology, 5: 471-478.

Shen, S. C. 1994. Mugilidae. Pp. 437-441. In: Shen, S. C. (ed.) Fishes of Taiwan. (In Chinese).

Smith, J. L. B. 1935. The fishes of the family Mugilidae in South Africa. Annals of the South African Museum, 30 (5): 587-644.

Smith, J. L. B. 1948. A generic revision of the mugilid fishes of South Africa. Annual Magazine of Natural History (South Africa), 11 (14): 833-843.

Smith, J. L. B. 1949. The sea fishes of southern Africa. Central News Agency, South Africa, 550 pp. Smith, M. M. 1975. Common and scientific names of the fishes from southern Africa. Special Publications of the J. L. B. Smith Institute of Ichthyology, 14 (1): 1-178.

Song, J, K. 1981. Chinese mugilid fishes and morphology of their cephalic lateral line canals. Sinozoologia, 1: 9-21.

Spain, A. V., Grant, C. J. & D. F. Sinclair. 1980. Phenotypic affinities of 11 species of Australian mullet (Pisces: Mugilidae). Australian Journal of Marine and Freshwater Research, 31: 69-83.

Starks, E. C. 1900. The osteological characters of the fishes of the suborder Percesoces. Proceedings of the United States National Museum, 22: 1-10.

Stead, D. G. 1908. Edible Fishes of New South Wales. Government Printing Service, Sydney, 123 pp.

Stiassny, M. L. J. 1990. Notes on the anatomy and relationships of the Bedotiid fishes of Madagascar, with a taxonomic revision of the genus Rheocles (Atherinomorpha: bedotiid). American Museum Novitates, 2979: 1-33.

Stiassny, M. L. J. & J. A. Moore. 1992. A review of the pelvic girdle of atherinomorph fishes. Zoological Journal of the Linnaean Society, 104: 209-242.

Stiassny,M. L. J. 1993.What are greymullets? Bulletin of Marine Science, 52 (1): 197-219.

Sulya, L. L., Box, B. E. & G. Gunther. 1960. Distribution of some blood constituents in fishes from the Gulf of Mexico. American Journal of Physiology, 199 (6): 1177-1180.

Svichenskaya, A. A. 1973. Mullet-like fossils in the USSR. Trudi Paleontologii Instituta Akademii Nauk (SSSR). Moscow, 138: 138:1-64. (In Russian).

Sunny, K. G. 1971. Morphology of the vertebral column of Mugil macrolepis (Smith). Bulletin of the Department of Marine Oceanography, University of Cochin, 5: 101-108.

Suvatti, C. 1981. Fishes of Thailand. Rungsilpa Ltd., Bangkok, 379 pp.

Thomson, J. M. 1954. The Mugilidae of Australia and adjacent seas. Australian Journal of Marine and Freshwater Research, 5: 70-131.

Thomson, J. M. 1963. Synopsis of Biological data on the grey mullet Mugil cephalus Linnaeus 1758. CSIRO Division of Fisheries and Oceanography, Fish Synopsis No. 1., Canberra, variable page numbers.

Thomson, J. M. 1964. A Bibliography of Systematic References to the Grey Mullets (Mugilidae). CSIRO Division of Fisheries and Oceanography, Technical Paper 16, Canberra, 127 pp.

Thomson, J. M. 1966. The grey Mullets. Annual Reviews of Oceangraphy and Marine Biology, 4: 301-335.

Thomson, J. M. 1974. Mugilidae (mullets). In: Fischer, W. and G. Bianchi (eds.) FAO Species Identification Sheets for Fishery Purposes. Fishing Area 57,71 (East Indian Ocean and West Central Pacific) Vol. 3. FAO, Rome, variable page numbers.

Thomson, J. M. 1975. The dentition of grey mullets (abstract). Aquaculture, 5: 108

Thomson, J. M. 1978. A field guide to the Common Sea and Estuary Fishes of non-tropical Australia. Collins, Sydney, 144 pp.

Thomson, J. M. 1980. Family Mugilidae. Grey Mullets. Pp 162-166. In: McDowall, R. M. (ed) Freshwater Fishes of South Eastern Australia (New South Wales, Victoria and Tasmania). Reed Pty. Ltd., Sydney, 208 pp.

Thomson, J. M. 1981. The taxonomy of grey mullets. In: Aquaculture of grey mullets (Ed. O.H. Oren) 1-15. Cambridge University Press, London.

Thomson, J. M. 1997. The Mugilidae of the world. Memoirs of the Queensland Museum, 41 (3): 457-562.

Tosh, J. R. 1903. Notes on the habits, development, etc., of the common food fishes of Moreton Bay. Queensland Harbour Marine Department Annual Report, Brisbane, pp. 17-24.

Trewavas, E. & S. E. Ingham. 1972. A key to the species of Mugilidae (Pisces) in the north-eastern Atlantic, with explanatory notes. Journal of Zoology, London, 167: 12-24.

Valenciennes, A. 1836. Mugiloides. In: Histoire Naturelle des Poissons (vol. 11) (Eds. G. Cuvier and A. Valenciennes,) 1-127. F. G. Levrault, Paris.

Van Der Elst, R. P. & J. H. Wallace. 1976. Identification of the juvenile mullet of the east coast of South Africa. Journal of Fish Biology, 9: 371-374.

Waite, E. R. 1904. Synopsis of the fishes of New South Wales. Memoirs of the New South Wales Naturalists’ Club, 2: 1-59.

Weber, M. 1894. Die Süsswasser-Fische des Indischen Archipels, Nebst Bemerkungen über den Ursprung der Fauna von Celebes. Zoologische Ergebnis Indien, 2: 405-476.

Weber, M. & L. F. de Beaufort. 1922. The Fishes of the Indo-Australian Archipelago (vol. 4). Leiden, 410 pp.

Whitehouse, R. H. 1927. The grey mullets of Tuticorin. Madras Fisheries Bulletin, 15 (3): 71-98.

Whitley, G. P. 1930. Five new generic names for Australian fishes. Australian Zoology, 6 (3): 250-251.

Whitley, G. P. 1941. Ichthyological notes and illustrations. Australian Zoology 10 (1): 1-52.

Whitley, G. P. 1945. New sharks and fishes fromWestern Australia. Part 2. Australian Zoology 11 (1): 1-42.

Whitley, G. P. 1948. New sharks and fishes from Western Australia. Part 4. Australian Zoology, 11 (3): 259-276

Wongratana, T., Senou. H. & C. Vittayanon. 1984. First record of the stripped grey mullet Mugil cephalus (Pisces: Mugilidae) from Thailand, with a key to all local species. Natural History Bulletin of the Siam Society, 32 (1): 11-20.

Yoshino, T. & H. Senou. 1984. Family Mugilidae. In: The Fishes of the Japanese Archipelago (Eds. H. Masuda, C. Amoaka, T. Araga, T. Uyeno and T. Yoshino) (English version). 119-121. Tokai University Press, Tokyo.  

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